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Research ArticleResearch Articles

Growth Rates of Mature Picea rubens, Abies fraseri, and Associated Hardwoods in Southern Appalachian Sky Islands

Rose C. Wetzel, David Carter, P. Corey Green and W. Mark Ford
Ecological Restoration, April 2026, 44 (1) 35-46; DOI: https://doi.org/10.3368/er.44.1.35
Rose C. Wetzel
Department of Forest Resources and Environmental Conservation, Virginia Polytechnic Institute and State University, 310 West Campus Drive, Blacksburg, VA, USA 24061,
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  • For correspondence: r.wetzel118{at}gmail.com
David Carter
Michigan State University, Lansing, MI, USA
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P. Corey Green
Department of Forest Resources and Environmental Conservation, Virginia Polytechnic Institute and State University, Blacksburg, VA, USA
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W. Mark Ford
U.S. Geological Survey, Virginia Cooperative Fish and Wildlife Research Unit, Blacksburg, VA, USA
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Abstract

Southern Appalachian spruce-fir sky islands are globally threatened, boreal-relict forests where Picea rubens (red spruce) and Abies fraseri (Fraser fir) are dominant. Stands are primarily composed of Fraser fir above 1,890 m with spruce-fir and spruce-dominated stands between 1,620 and 1,890 m and hardwood associates below 1,620 m. Hardwood encroachment and the reduction of spruce-fir forest cover are primary concerns as climate change increases droughts and raises temperatures, threatening high elevation-dependent systems. We compared growth rates (basal area increment; cm2 yr-1) of canopy red spruce, Fraser fir, and competing hardwoods among forest cover types and topographic aspects at six southern Appalachian sky islands in North Carolina, Tennessee and Virginia. We expected slower growth of red spruce and Fraser fir than hardwoods at ecotonal transitions, lower elevations, and on more xeric aspects. However, we observed that red spruce and Fraser fir growth rates were similar to or greater than those of hardwoods. Specifically, red spruce growth rates were greater than those of hardwoods on northern and southern aspects in the ecotonal transition whereas Fraser fir growth rates were often greatest in fir-dominated stands. Our findings could assist managers in prioritizing restoration treatment locations to expand the spruce-fir cover type downslope in the southern Appalachians.

Keywords
  • basal area increment
  • boreal-relict forests
  • climate change
  • cold-adapted conifers
  • tree growth
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Ecological Restoration: 44 (1)
Ecological Restoration
Vol. 44, Issue 1
1 Apr 2026
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Growth Rates of Mature Picea rubens, Abies fraseri, and Associated Hardwoods in Southern Appalachian Sky Islands
Rose C. Wetzel, David Carter, P. Corey Green, W. Mark Ford
Ecological Restoration Apr 2026, 44 (1) 35-46; DOI: 10.3368/er.44.1.35

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Growth Rates of Mature Picea rubens, Abies fraseri, and Associated Hardwoods in Southern Appalachian Sky Islands
Rose C. Wetzel, David Carter, P. Corey Green, W. Mark Ford
Ecological Restoration Apr 2026, 44 (1) 35-46; DOI: 10.3368/er.44.1.35
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Keywords

  • basal area increment
  • boreal-relict forests
  • climate change
  • cold-adapted conifers
  • tree growth
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