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Department of Forest Resources and Environmental Conservation, Virginia Polytechnic Institute and State University, 310 West Campus Drive, Blacksburg, VA, USA 24061,

Michigan State University, Lansing, MI, USA

Department of Forest Resources and Environmental Conservation, Virginia Polytechnic Institute and State University, Blacksburg, VA, USA

U.S. Geological Survey, Virginia Cooperative Fish and Wildlife Research Unit, Blacksburg, VA, USA